Potting Media for Plumeria

Low-quality or substandard potting media is a major source of problems with plumeria grown in containers (and, really, any container-grown plants). Plumerias that are potted in substandard or old decomposed potting soil are weaker than healthy plants, so they’re more likely to drop leaves, turn yellow, attract pests and just generally die.

Potting media, of course, is just one of the factors that contribute to healthy plumeria. But because most people only re-pot  when they have too, it’s actually the hardest single factor to quickly control. Other factors that are much easier to control include moisture levels, pH, humidity, temperature, and light levels.

Because potting soil is so important and relatively hard to change, it’s extra important to pay careful attention to your potting media and start off with the best possible mix. Your plumeria will thank you for it, and ultimately, you’ll end up with healthier plumeria and more blooms.

A good potting media should:

  • Provide support for the growing plant
  • Provide adequate drainage, very important for Plumeria
  • Provide adequate available nutrients (usually through a fertilizer and/or organic matter added to the soil)
  • Provide adequate aeration around the roots

You can easily find many good potting mixes. Most are based on sphagnum peat moss, with additives like compost, humus, perlite, vermiculite and pumice. Some have water-retention crystals, these are not good for plumeria. You can also find soils with fertilizer added, pine bark, and other ingredients like coir, seaweed and worm castings. I prefer to use a non fertilized soil mix, that way I know what I’m starting with.

For plumeria it is important the soil you select allows for adequate drainage, it has air pockets around the root zone and critically, and how long does it last before the inevitable decay sets in (all organic things decay) and starts to lose its desirable qualities?

Using Soil and Soil Mixes

Well prepared garden soil is great for growing Plumeria in the ground but when it comes to growing Plumeria in containers, it needs to be changed improved or changed as the roots grow and use up the nutrients.

Soils for containers need to be well aerated and well drained while still being able to retain enough moisture for plant growth.

When choosing what to use to fill containers, never use garden soil by itself no matter how good it looks or how well things grow in it out in the garden. When put into a container both drainage and aeration are severely impeded, and the results are that plumeria grows poorly or not at all.

Soils for containers need to be modified in some way to ensure proper drainage and aeration. Container soils are often referred to as soilless or artificial media because they contain no soil at all. They are often composed of various things such as peat, perlite, vermiculite, bark, coir fiber (ground coconut hulls) in a variety of recipes depending on the manufacture and the type of plant material being grown. They can be found under a variety of trade names and in sizes ranging from a few quarts to bales that are many cubic feet in size. The choice of growing media for plumeria are well drained and don’t retain a lot of moisture over a long period of time. You should choose media that’s courser in texture containing more bark, perlite or sand.

When soil mixes are used, they should be moistened slightly before planting. Fill a tub with the media, add water and lightly fluff the media to dampen it.

Some garden soils can be used as a container media but it needs to be modified or amended. An acceptable soil based mix can be made by using one part garden soil, one part peat moss and one part perlite or coarse builders sand. Don’t use fine beach sand or play sand.

Advantages and Disadvantages

The disadvantage is that the soil you are using may contain insects, weed seeds and disease organisms. Soilless media are generally free of these things. Soil-based media are heavier. This may be an advantage though especially if the containers are used in a very windy location and the extra weight will help keep the pot upright. Soil-based media are also a bit more forgiving when it comes to water and fertility. They tend not to dry out as fast, and they also tend to hold on to nutrients longer. Because soilless media can be a bit more costly, you could mix 25 percent soil with the soilless media to stretch your soilless media for a few more pots.

Can soilless media be reused from year to year? If the plants in the containers were healthy during the growing season with no major disease issues, you could remove the plant material at the end of the season and reuse the media next season. A good way to make sure you soil disease free by putting the soil in a black plastic bag and keep in full sun for a week or so, rotating the bag every few days. You may also have to add some additional new media to fill the container. There will come a time when the media in the container will need to be replaced. Over the course of time, the organic materials that the soilless media is made from a breakdown and decompose to the point where you will lose a lot of nutrients, drainage and aeration properties that are inherent in soilless container media. When that happens, discard the media to the compost pile or to the garden and refill the container with fresh media.

There are also bagged products labeled as topsoil that tend to be largely sedge peat. While they are inexpensive and look very good, once put into a pot they are poorly drained and poorly aerated. They can be used in combination with soilless media products or amended much the same way as if you were using garden soil.

When filling containers with media, don’t fill the pot to the top. Leave about a one-inch space between the top of the soil and rim of the pot. This will help make watering the pot easier as it provides a place to “put water” and not have it run over the edge.

Filling very large containers can be costly especially when using commercially prepared media. To reduce the cost and also the weight of the container consider adding “filler” to the bottom of the container to take up space. Many things can be used, but chipped mulch will decompose and add nutrients to your pot over the course of the growing season.

Guide to Coco Coir Growing Media

When picking out the right growing media for Plumeria there are several things to consider.  After reading the labels, you actually became more confused than before.

The scoop on growing media. First off, it needs to be stated that technically all of the “potting soils” sold in grow shops and gardens centers are actually soilless mixes because they do not contain any of the natural soil components including: sand, silt or clay. Choosing your media, watering and fertilizing habits are key to successful plumeria growing.

Soilless Growing Media

Soilless growing is great and has easily become the method of choice for plumeria growers because of its superior properties, and ease of amending. Soilless media allows for the ultimate control over desired inputs, drainage properties, pH and nutrient control. The most common soilless potting mixes are made with one of two bases.

The most common base is Coco Coir. This media is 100% natural and made from coconut husk fibers. It is generally a finely ground blend with some desired variation. It has the perfect air to water ratio, and thus is hard to over water. Coco coir also has a high CEC encouraging optimal nutrient release. Due to its superior qualities Coco coir allows for rapid growth. Keep in mind that with Coco coir bases, Calcium -Magnesium supplementation may be needed.

If it is not a coco coir base, it is generally a peat based mix. Peat is partially decayed and dried sphagnum moss. It is extremely high in organic matter, and retains moisture better than coco coir. Thus, it is not as forgiving if you over water. By itself, peat usually has a very low pH, between 3 and 5. In potting mixes you can help balance the acidity with the addition of lime. For a peat based potting mix our go to of choice is Pro-Mix.


Once you’ve decided which base is best for your growing conditions, next you have the soilless amendments. Amendments help define the final properties of the mix, which is customized for health growth results. Common amendments include the following, and added for the following reasons.

  • Perlite: increases drainage and aeration
  • Dolomite lime: buffers pH, provides calcium and magnesium
  • Worm castings: natural source of nitrogen, enhances beneficial microbe population
  • Mycorrhizae: symbiotic growth on and around roots, increases water and nutrient uptake
  • Azomite: buffers pH, slow release of micronutrients
  • Oyster shell: buffers pH, provides calcium
  • Dried kelp: source of potassium and natural plant hormones
  • Alfalfa meal: organic source of balanced fertilizer, amino acids and triacontanol
  • Feather meal: source of organic slow release nitrogen
  • Fishbone meal: organic source of phosphorus and calcium

Typical Ground or Garden Soil

Soil is used almost exclusively in outdoor growing. Its composition varies from place to place and consists of countless natural and native components. These components include mineral particles- sand, silt, and clay (45% collectively), air (20-30%), water (20-30%), organic matter (1-6%), microorganisms and the ever pervasive weed seed.  Organic (carbonaceous) matter, often referred to as humus, is composed of decaying plant and animal remains. Microorganisms include both beneficial and harmful bacteria and fungi. Weed seeds are one of the grower’s greatest foes. Not only are they a nuisance, but they also compete with your crop for water and nutrients.

Since the composition changes from area to area, the properties change along with it. Soil with a healthy ratio of soil particles and organic matter will have good drainage. If the soil is compacted with low aeration, roots and plant growth will struggle. Conversely, if there is too much drainage, growth will also be hindered due to a lack of water and nutrient retention.

Along with your soil composition changing from area to area with soil, so will your soil fertility. The most important aspects of soil fertility to keep in mind include the presence and amount of organic matter, macro and micro nutrients, pH and CEC. There are various amendments and fertilizers that can help you achieve desired results in an outdoor setting.

Soil pH and its Importance

Soil pH

(a measure of the acidity or alkalinity of the soil) Soil pH is one of the most important soil properties that affects the availability of nutrients.

  • Macronutrients tend to be less available in soils with low pH.
  • Micronutrients tend to be less available in soils with high pH.

Lime – can be added to the soil to make it less sour (acid) and also supplies calcium and magnesium for plants to use. Lime also raises the pH to the desired range of 6.0 to 6.5.

In this pH range, nutrients are more readily available to plants, and microbial populations in the soil increase. Microbes convert nitrogen and sulfur to forms that plants can use. Lime also enhances the physical properties of the soil that promote water and air movement.

It is a good idea to have your soil tested. If you do, you will get a report that explains how much lime and fertilizer your crop needs.


pH Testing

Because of the specific requirements for proper mineral uptake it is important to test the pH of three things: your water, your fertilizer solution and your growing medium.  Litmus paper, and pH drops are reliable enough for generalizations and maintaining a healthy system.  However, if a problem is noticed, then a more accurate method should be employed.  A high quality pH meter can be used to test water, fertilizer solution, as well as growth medium. The most popular tool used to test pH is BlueLab pH Pen because of its versatility and ease of use. We also suggest the Myron L Ultrapen PT2 for the most accurate reading. Unlike its competitors it reads the pH to the hundredths place, which can be very important when you consider that pH is a logarithmic scale. For a more in depth look at your solution’s total dissolved solids, we suggest the BlueLab Combo Meter. This meter measures pH, EC, has a replaceable probe and a 5 year warranty.

pH Adjusting

In fertilizer solutions, using a pH up or down solution may be required in order to achieve the preferred acidity or alkalinity. PH up generally consists of Potassium Bicarbonate and/or Sodium Bicarbonate.  PH down is most popularly Phosphoric Acid and/or Citric Acid.  The ingredients of the pH solution used, as well as the ingredients of the nutrient solution will determine short term and long term stability of the pH of the solution.  Bicarbonates can have long lasting effects on the media environment and should be used in hydroponic gardening, or very sparingly for soil.  In soil, Dolomite Lime should be the go-to ingredient for long term pH stabilization.  Phosphoric Acid, which is ideal for hydroponics, can keep solutions stable for much longer than a Citric Acid based pH solution, but may not be better to acidify a soil grow. One of our favorite organic pH adjusters is Vitazyme and Carl Pool’s Root Activator.

Perlite vs Vermiculite

Perlite and vermiculite are both used to improve moisture retention and aeration in soil. They are used in a similar manner, but they are not interchangeable. Perlite and vermiculite are quite different in composition and in how they improve your soil. Determining which is better for use in your garden depends on your plants and their needs.


Vermiculite is an aluminum-iron-magnesium silicate that resembles mica in appearance. For use in horticultural applications, vermiculite is heated to expand the particles. This expansion enables it to absorb moisture when used as a potting medium. Vermiculite can soak up 3 to 4 times its volume in water. It also attracts plant nutrients such as potassium, magnesium, calcium and phosphorus. Perlite is an amorphous volcanic rock that is rich in silicon. When mined for use as a potting medium, perlite is crushed and also heated to expand the particles. The microscopic bubbles in perlite granules absorb and hold water as well, but they also hold air.


Vermiculite is a spongy material that is dark brown to golden brown in color. It is shaped like flakes when dry. Perlite is a porous pumice-like material that looks like white granules. Sometimes perlite is mistaken for tiny plastic foam balls when used in potting soil mixtures.

Water Retention

Perlite and vermiculite are both good at retaining water, but vermiculite acts more like a sponge, holding much more water than perlite and offering less aeration for the plant roots. Perlite retains water because of its large surface area with nooks and crannies available for water storage. Because it is porous it allows excess water to drain more readily than vermiculite and improves soil aeration.

Horticultural Uses

Both vermiculite and perlite are used in the garden to prevent soil compaction, improve aeration and retain moisture. They’re also used in propagation of new plants and seed cultivation, as well as in indoor container growing, composting and on lawns. However, the way that each material retains water, and how much water is retained, makes each one suitable for different plants. Vermiculite is ideal for plants that prefer lots of water, such as forget-me-nots and some irises. Perlite would dry out too rapidly for water-loving plants. However, the amount of water vermiculite holds is not ideal for plants such as cacti or rhododendrons, which need a well-drained soil. The moisture retained by vermiculite would lead to root rots or plant death.

Perlite vs Vermiculite: What’s the Difference?

The difference between perlite vs vermiculite is important to know for the prosperity of your garden. They seem very similar, but differ in a few crucial ways.

What is Perlite?



Perlite is lightweight, easy to handle, clean and has no odor. It has a pH of 6.6 to 7.5.

The life of a bag of perlite begins as volcanic glass — but not any type of volcanic glass. It’s formed when obsidian contacts water, creating a unique type of volcanic glass with a high water content. When manufacturers apply heat to perlite, it puffs up into little white balls. Often times they’ll mix these little white balls — what we call perlite — into potting soils to aid with soil aeration and water retention. It retains some water but also air on the surface of the little balls in all the hidden nooks and crannies.

Perlite is a good choice when you have plants in your garden that require soil to dry out completely between watering. For example, if you’re growing a cactus or a succulent, perlite is a great addition to the soil.

Because it’s so porous, perlite does allow excess water to drain quickly…sometimes all over your porch. It has a tendency to easily crush into a powder between your fingers, but this usually isn’t a problem because it doesn’t encounter that type of pressure in your pots or beds. It’s chief use is to improve soil aeration, lightening the soil and giving better drainage and oxygen access for your plants’ roots.

What is Vermiculite?



Vermiculite interacts with potassium, calcium and magnesium in your soil. It also helps to raise the pH slightly of your plants even though it’s a neutral pH of 7.0.

Vermiculite is made from compressed dry flakes of a silicate material which is absorptive and spongy. The color of vermiculite is a golden brown to a dark brown and is a sometimes difficult to tell from the potting soil it’s mixed with. When water is added to vermiculite, the flakes expand into a worm-like shape and act like an absorbing sponge. If you want to poke these “vermiculite worms” with your fingers, you’re not alone — that’s what I wanted to do when I first saw them too!

Vermiculite is best used for plants that require soil to stay damp and not dry out. For plants that love water, using vermiculite or mixing a healthy scoop of it into your potting soil is the way to go. It can absorb 3 to 4 times its volume when water is added, making your pots a little bit on the heavy side.

Since vermiculite acts like a sponge and absorbs more water than perlite, it doesn’t aerate the soil as well. This means less oxygen for plant roots. If you use it when growing plants that don’t need damp soil, you might find your plants suffering from root rot. So be aware of your plants’ needs when you decide how water retentive you want your soil to be.

More Differences Between Vermiculite and Perlite

There are major differences between vermiculite and perlite, making it important to choose the right one, lest your garden be ruined by a bad growing media choice.

We’ve already covered the biggest difference: Vermiculite will mix with soil and help to retain water. Perlite, on the other hand, will add drainage to the soil that it’s mixed with.

Vermiculite finds its way into many seed starting systems. It both protects seedlings from fungus that so often ruins seed starting, and helps to retain water in the tiny little pods that seeds start in. While perlite can be used with seedlings, it’s better used when you move your seedlings into separate pots for additional drainage.

Which To Use In Your Garden?

There’s a large discussion in the gardening community on which to use in the garden. Here’s the truth: it’s a false debate. They both have their own purposes in the garden.

Use Perlite If…

  1. You have plants that need to dry out before watering again
  2. When you move your seedlings to separate pots
  3. You need to loosen clay soil in your garden

Perlite when added to clay soils, it can eliminate both surface crusting and puddles. It will also help to reduce fluctuations in soil temperatures in your garden soil. Perlite will also improve both drainage and aeration in your home gardens. Horticultural perlite can be bought in different grades according to how you’re going to use it. For general application, a fine to medium grade can be used. It’s free of weeds, disease free and sterile.

Use Vermiculite If…

  1. You need an additive for plants that need to be kept moist
  2. You want your seed trays to develop strong seedlings

Vermiculite is odorless, can be purchased in horticultural-grade bags with directions on working it into the garden soil. It’s a permanent soil conditioner and won’t break down in your soil like compost does. When it is watered or it rains, the vermiculite will hold water in the soil until the soil begins to dry out and releases it. Vermiculite can be used in potted containers, on lawns and for composting. It can be used in mycology for mushrooms added to the substrate.  It can improve the soil that needs an additive to retain water for your plants which need it.

In summary: Both are good additives to your gardening needs, you just need to know what you’re using them for!

Soils and Growing Mediums

What Is Soil?

Soil is a living, breathing, natural entity composed of solids, liquids, and gases.

Soil has five major functions:

  1. Provides a habitat for organisms
  2. Recycles waste products
  3. Filters water
  4. Serves as an engineering material
  5. Provides a medium for plumeria growth

Our focus will be on the fifth function. In this role, soil provides structural stability for plumeria and retains and relinquishes water and the nutrients necessary for plumeria growth.

An ideal soil for plant growth contains 50% porespace and 50% solids, with the porespace filled with equal parts air and water. This distribution rarely occurs because porespace varies with soil texture and soil management. For example, tilling increases porespace, while poor drainage and compaction reduce it.

Soil solids are a blend of mineral materials and organic matter. The mineral materials are typically weathered rock of varying sizes called sand, silt, and clay. The organic matter consists of decaying plant and microbial residues. The relative amounts of porespace and mineral and organic matter vary greatly among different soil types. But for plumeria growth, most soil scientists agree that 50% porespace, 45% mineral matter, and 5% organic matter make up an ideal ratio. The distribution of soils and porespace in compacted in poorly drained soil.

Even a small amount of organic matter can have a dramatic effect on the physical, chemical, and biological properties of soil.

1Brady, N. C. and R. R. Weil. 2004. Elements of the Nature and Properties of Soils, 2nd Edition. Atlanta, GA: Prentice Hall.

The Soil Profile

Most naturally occurring, undisturbed soils have three distinct layers of variable thicknesses. The layers are the topsoil, subsoil, and parent material. Each layer can have two or more sublayers called horizons. Collectively, the horizons make up the soil profile. The predominate parent material varies by location.

Soils’ properties vary with the soil depth. The surface soil, or topsoil layer (O and A horizon in Figure 1–2), usually contains less clay, but more organic matter and air, than the lower soil layers. Topsoil is usually more fertile than the other layers and has the greatest concentration of plant roots.

The subsurface layer (B and C horizon in Figure 1–2), known as subsoil, usually has a higher clay content and lower organic matter content than the topsoil.

Soil properties often limit the depth to which plant roots can penetrate. For example, roots will not grow through an impenetrable layer. That layer may be bedrock (Figure 1–3), compacted soil, or a chemical barrier, such as an acidic (very low) pH. A high water table can also restrict root growth due to poor soil aeration. Few big trees grow in shallow soils because big trees are unable to develop a root system strong enough to prevent them from toppling over. Shallow soils also tend to be more drought-prone because they hold less water and thus dry out faster than deeper soils. Water lost to runoff on shallow soils would instead be absorbed by a deeper soil. In addition, deep soils allow the roots to explore a greater volume, which means the roots can retain more water and plant nutrients.

Soils change in three dimensions. The first dimension is from the top to the bottom of the soil profile. The other two dimensions are north to south and east to west. The practical meaning of this three-dimensional variability is that as you move across a state, a county, or even a field, the soils change.

Five factors of soil formation account for this variation:

  1. Parent material
  2. Biological activity
  3. Climate
  4. Topography
  5. Time

Differences in even one of these factors will result in a different soil type. Soils forming from different parent materials differ. Soils forming from the same parent material in varying climates differ. Soils at the top of a hill differ from soils at the bottom. The top of the hill loses material due to natural erosion; the bottom gains the material from above. Considering the number of possible combinations of these five factors, it is not surprising that more than 450 unique soil series are currently mapped in North Carolina. Globally, more than 20,000 different soil series occur.



Physical Properties of Soil

The physical properties of soil are characteristics that can be seen, felt, or measured. These include color, texture, structure, and water-holding capacity. Such properties usually determine the suitability of soil as a growth medium. Some physical properties, such as texture, are not economically feasible to change on a large scale.

A soil’s fertility, which is a chemical property, is easier to change than the soil’s physical properties.


Organic matter, the soil minerals present, and the drainage conditions all influence soil color. Color alone is not an indicator of soil quality, but color does provide clues about certain conditions. For example, light or pale colors in grainy topsoil are frequently associated with low organic matter content, high sand content, and excessive leaching. Dark soil colors may result from poor drainage or high organic matter content. Shades of red indicate a clay soil is well-aerated, while shades of gray indicate inadequate drainage (Figure 1–4). In well-drained soils of the NC mountains and piedmont, the subsoil colors are often shades of red, brown, and yellow. In poorly drained soils, the subsoil is grayer in color.


Soil texture, which refers to the proportions of sand, silt, and clay, influences nearly every aspect of soil use and management. Sand is the largest particle (at 2.0 to 0.05 mm), silt is much smaller (0.05 to 0.002 mm), and clay is the smallest (less than 0.002 mm) (Figure 1–5). To compare particle sizes, imagine that a sand particle is the size of a basketball. On that scale, a silt particle would be the size of a marble, and a particle of clay would be a pinpoint. How fine (clayey) or coarse (sandy) a soil is will determine many of the soil’s physical and chemical properties.

Much of a soil particle’s ability to react with water and nutrients is related to the amount of surface area available. When the individual particle size is small, more individual particles will fit in a given space, and thus make more surface area available. Clay, with its tiny particle size and platelike structure, holds water and nutrients effectively, while sand, which has a large chunky structure, does not. In addition to being smaller, clay particles are composed of different minerals than sand and silt, and a clay particle’s structure is more like a stack of paper plates than a grain of sand (Figure 1–6).

Table 11. Particle type, number of particles per gram, and the average surface area per gram.

Particle TypeDiameter (mm)Number of Particles per gramSpecific surface area (cm2/g)
Clay< 0.00290,260,853,0008,000,000
Coarse sand1.00-0.5072023
Fine sand0.25-0.1046,00091
Medium sand0.50-0.255,70045
Very coarse sand2.00-1.009011
Very fine sand0.10-0.05722,000227

Rocks and Gravel

Rocks and gravel, which are large, coarse materials, can be found in many soils, but they are not considered when determining soil texture. Although some rocks and gravel in the soil will not affect plant nutrient uptake, they can make the soil difficult to dig. If the garden is mostly rocks or gravel, the soil will have a reduced water- and nutrient-holding capacity, and will be unfit for growing plants. In such a situation, it may be easiest to install raised beds and import soil.

The relative proportions of sand, silt, and clay determine a soil’s textural class (Figure 1–7). For example, a soil that is 12% sand, 55% clay, and 33% silt is in the clay textural class. Soil texture is a permanent feature, not easily changed by human activity. Consider a typical mineral soil that is 6 inches deep on 1 acre. That soil weighs about 2 million pounds. To change the sand content just 1% would require adding 20,000 pounds (or 10 tons) of sand. A 1% change in sand content would have minimal effect. A significant effect might require a 10% change, which would mean adding 100 tons of sand.

Adding organic matter is a more economically feasible alternative for improving soil. Adding organic matter does not change a soil’s texture—the percentage of sand, silt, and clay in the soil—but adding organic matter will alter soil structure by increasing the porespace and improving drainage. Gardeners can be successful with any soil texture, as long as they know the attributes and limitations of that soil.

Typically, laboratory procedures are used to determine the soil texture. It is possible, however, to use the procedure outlined in Figure 1–8 to determine the textural class by the “feel” method. It takes practice and calibration, but it can provide a reasonable estimate of the soil texture.

Sandy or Coarsely Textured Soils (Figure 1–9)

  • Low in organic matter content and native fertility.
  • Rapidly permeable and do not hold soil moisture.
  • Nutrient leaching is a concern, so proper fertilization is a must. Apply smaller amounts of nutrients, and apply them more frequently.
  • Low in cation exchange and buffer capacities.
  • Well-suited for road foundations and building sites.
  • Feel gritty.

Loamy or Medium-Textured Soils (Figure 1–10)

  • Contains more organic matter.
  • Permit slower movement of water and are better able to retain moisture and nutrients.
  • Are generally more fertile.
  • Have higher cation exchange and buffer capacities.
  • Feel crumbly.

Clayey or Finely Textured Soils (Figure 1–11)

  • Higher nutrient-holding capacity.
  • Higher available water-holding capacity.
  • Finely textured soils exhibit properties that are somewhat difficult to manage or overcome.
  • Often too sticky when wet and too hard when dry to cultivate.
  • May have shrink-and-swell characteristics that affect construction uses.
  • Feel slippery.

How Do Soil Types Affect Gardeners?

Compaction. Compaction occurs when pressure is applied to soil particles and the air and water are pushed out of the porespaces. Large, cubic sand particles are not easily compacted. Clay particles, small and platelike, are easily aligned and can compact, especially when wet. Compaction inhibits the movement of water, gases (air), and roots. Compacted soils have less infiltration, greater runoff, a higher risk of erosion, and more restricted root growth than soils without compaction. Water drains slowly, which may increase the likelihood of plant root diseases.

Erosion. Sand particles are heavy, so they are not easily picked up and moved by water or wind. Clay particles are sticky, so they are not easily moved. Silty loam particles are light and not sticky, so erosive forces easily move them. Eroded soils are usually harder to till and have lower productivity than soils without erosion. The main causes of soil erosion in North Carolina are insufficient vegetative or mulch cover, and improper equipment and methods used to prepare and till the soil (Figure 1–12).

Soil erosion can be minimized by following a few preventive measures:

  • Choose plants suited to the soil so they establish well.
  • Mulch the surface each year with organic materials 1 inch to 3 inches deep.
  • Adequately fertilize to promote vigorous, but not excessive, plant growth.
  • Create a water diversion, such as a grass waterway, to capture and slow water movement.
  • Align rows to follow the land’s contour so that water flowing downhill is slowed.
  • Use proper tillage methods, such as not tilling when the soil is overly wet and not overtilling.
  • Plant a winter cover crop.
  • Consider installing rain gardens to capture sediment and runoff.

Surface Area. The most active part of a soil particle is its surface area. A particle’s surface is where nutrient exchange takes place. Sand particles have a small surface area relative to their mass, meaning they do not hold onto nutrients well. Clay particles have a large surface area relative to their mass, so a small amount of clay can add a significant amount of surface area to a soil, increasing the nutrient-holding capacity.


Soil structure refers to the grouping of individual soil particles into larger pieces called peds or aggregates. The structure of topsoil is usually granular and resembles chocolate cookie crumbs (Figure 1–13). Good granular structure allows rapid movement of air and water within the soil. Poor granular structure decreases movement of air and water. Good soil structure allows for extensive root development; poor structure can limit root growth. Supplying an adequate amount of organic matter and working the soil only when it is not excessively wet promotes good topsoil structure.

Water-Holding Capacity

Water enters the soil from precipitation or irrigation. It exits by draining from the soil, evaporating from the surface, and through transpiration from plant leaves. Water-holding capacity—the retention of water moving through soil—depends on differences in soil porespace. Ideal soils are half porespace with equal amounts of air and water filling the pores. Too much air means plants will wilt. Too much water means reduced plant vigor and susceptibility to root rot, which occurs due to anaerobic conditions.

Soils differ in the number of large (macro), medium (meso), and small (micro) pores. Macropores, which are more common in sandy soils, take up water more quickly and drain faster than meso- and micropores. This rapid draining from macropores is called “gravitational water” because the weaker forces of adhesion and cohesion in macropores cannot overcome gravity’s pull. Within 24 hours after a saturating rain, gravitational water reaches the lower soil horizons, and the soil is at field capacity: the meso- and micropores are still full of water because their adhesive and cohesive forces are stronger than gravity. Water in the mesopores is available to plants. But when the mesopores lose water as the soil dries through plant uptake and transpiration, soil moisture reaches the permanent wilting point. At the permanent wilting point, micropores are still full of water, but this water is so tightly held that it is not plant-available. Note that plants may wilt before the permanent wilting point if the plant transpires water through the leaves faster than it can take water up from the soil through its roots. This is why plants may wilt on hot days and then recover once the sun goes down and why plants can balance uptake with transpiration (Figure 1–14).

How to Remediate Compaction

Compaction is a likely problem if there has been recent construction or other traffic over the area. Deep cultivation, which is mixing the top 6 inches to 2 feet of soil with a tiller, disk, or hand tools, may be needed to loosen the soil. Incorporation of organic matter during deep cultivation can help to rehabilitate soil structure by creating aggregates and both macropores (for drainage) and mesopores (for plant-available water). Digging or cultivating soil when it is wet or excessively dry can destroy structure.

Be wary of quick fixes, such as starting over with a truckload of topsoil. Unfortunately, there are no standards on material sold as “topsoil.” New problems may be brought on site, such as weed seeds and disease organisms. Adding new topsoil to existing soil may also create drainage problems when water moves through the purchased topsoil and reaches the compacted layer. The water can pool and create unfavorable conditions for root growth.

Clay soils, which tend to hold excessive amounts of water and become compacted easily, present some tricky problems. Common mistakes are adding sand or peat moss to improve drainage. Adding sand to clay will reduce soil structure, lowering porespace. Adding peat moss will increase the clay soil’s high moisture-holding capacity. The best advice is to add smaller amounts of organic matter consistently every year, minimize compaction, and let soil biology naturally improve the structure over time.

Organic Matter

Organic matter consists of the remains of plants and animals and gives soil a gray to very-dark-brown color. Organic matter is home to many soil organisms.

Earthworms, insects, bacteria, fungi, and animals use organic matter as food, breaking it down to obtain energy and essential nutrients. Humus is the portion of organic matter that remains after most decomposition has taken place (Figure 1–16).

When organic matter decomposes in the soil, carbon dioxide is released and replaces some of the oxygen in soil pores. Carbon dioxide is dissolved by water in soil to form a weak acid. This solution reacts with soil minerals to release nutrients that can be taken up by plants. The digested and decomposing organic matter also helps develop good air-water relationships. In sandy soil, organic material occupies some of the space between the sand grains. This binds them together and increases water-holding capacity. In a finely textured or clay soil, organic material creates aggregates of soil particles. This allows water to move more rapidly around soil particles.

The amount of organic matter in the soil depends primarily on rainfall, air temperature, the kinds of plants that have been growing in a soil, management practices, soil temperature, and drainage. Soils that are tilled frequently are usually low in organic matter because tilling decreases residue particle size and increases the amount of air in the soil, increasing the rate of organic matter decomposition. Poorly drained soils tend to have a high percentage of organic matter because low oxygen levels limit decomposition organisms. To build organic matter in garden soil, till in compost when the garden is first created, but do not till in subsequent years. Instead, apply thin layers (1 inch to 3 inches), of organic mulch or compost to the soil surface each year (Figure 1–17). This material will break down, and the organic matter levels in the soil will gradually increase.

Improving the Soil

Good aeration and drainage, as well as the ability to hold adequate moisture and nutrients, are key components of an ideal soil environment. Although there is no cookbook recipe for creating this ideal environment, these are some of the most important strategies for improving soil quality:

  • Minimize soil compaction (do not walk on garden beds or work wet soil) (Figure 1–18).
  • Reduce drainage problems.
  • Decrease erosion.
  • Plant a cover crop (Figure 1–19).
  • Incorporate organic matter.
  • Provide a 1- to 3-inch layer of organic mulch on the soil’s surface.

Organic amendments can improve soils that suffer from high compaction, poor drainage, and erosion. Materials such as compost, manures, and pine bark are more effective and economical than vermiculite, peat moss, sand, topsoil, or perlite. Table 1–2 reviews the amounts of organic material to be added to soil per 100 square feet. When working in small areas, a general rule of thumb is to incorporate a 3- to 6-inch layer of organic material into the soil. The organic matter must be decomposed before plants can use the nutrients. The rate of decomposition of organic matter by soil organisms is affected by moisture, temperature, particle size, the carbon-to-nitrogen ratio, and nitrogen availability. The proper balance of carbon and nitrogen is needed for rapid decomposition, as are warm temperatures and adequate moisture. When using straw, leaves, or sawdust (which are high in carbon), add nitrogen fertilizer while the material is decomposing. Soil microbes use nitrogen during decomposition and may deprive plants, resulting in slow or stunted plant growth. Incorporating organic matter some months before planting the garden allows the material time to decompose and have plant-available nutrients in place for good plant growth.

Table 1–2. Organic Materials and Their Application Rates

Organic MaterialAmount to Be Added per 100 Square Feet
Compost10–20 cubic feet
Corncobs50 pounds (2 bushels)
Hay60 pounds (1 bale)
Leaves75 pounds (3–4 bushels)
Sawdust50 pounds (2 bushels)
Straw60 pounds (1 bale)
Wood chips50 pounds (2 bushels)

Incorporating Soil Amendments

Conditioning soil requires increasing organic matter content to 25% by volume. Incorporating a minimum of 2 inches of material into the top 6 inches of soil will create approximately 8 inches of amended soil. These additions raise the planting bed, improving drainage and making plants more visible. Incorporating more than 50% organic matter may negatively affect plant growth. Be careful when using organic material, making certain that it is fully composted and not merely aged. Microbes attracted to partially decomposed materials will compete with plants for nutrients, especially nitrogen and sulfur, resulting in nutrient deficiencies and poor plant growth.

The best organic matter amendments for clay soils are pine bark (less than 1/2 inch in diameter) and composted leaf mold. The following amendments are not recommended because they do not adequately improve the physical properties of clay soil: peat moss, sand, hardwood bark, wood chips, and pine straw.

For sandy soils, organic matter amendments, such as pine bark or compost, will improve water retention.

Figure 1–19. A cover crop of white rye grass was planted in this annual flower bed. It is being turned under to add nutrients to the soil before planting.

Chemical Properties of Soil

There are strong relationships between soil physical properties and soil chemical properties. For example, surface area is directly related to chemical reactivity.

Cation Exchange Capacity (CEC)

The negative ends of two magnets repel each other. The negative end of one magnet attracts the positive end of another magnet. This same principle affects the retention of plant nutrients in soil. Some plant nutrients are cations, which have a positive charge, and some are anions, which have a negative charge. Just like the opposite poles on magnets, cations will be attracted to anions.

Soil particles are similar to a magnet, attracting and retaining oppositely charged ions and holding them against the downward movement of water through the soil profile. The nutrients held by the soil in this manner are called “exchangeable cations” and can be displaced or exchanged only by other cations that take their place. Thus, the negative charge on a soil is called the cation exchange capacity (CEC). Soils with high CEC not only hold more nutrients, they are better able to buffer or avoid rapid changes in the soil solution levels of these nutrients. A soil test will tell you the CEC number of your soil. Soils high in clay, silt, or organic matter will have a CEC number of 10 or greater, and no remediation is needed. Sandy soils will have a CEC number between 1 and 5. Adding organic matter to these soils will help increase the CEC.

Too Much of a Good Thing: Nitrogen Leaching

Soil testing provides valuable information on pH and plant-available nutrients. Test your soil before planting and every two to three years thereafter. Inexpensive soil test kits are unreliable. To accurately determine your soil characteristics and the proper amount of lime and fertilizer to apply, contact the NC Department of Agriculture and Consumer Services (NCDA&CS). The accuracy of these reports, however, depends on the quality of the sample submitted.

Soil Testing Just like magnets, negative charges repel negative charges. Soils with high CEC tend not to hold anions. As a result, water moving through the soil profile will leach negatively charged nutrients, such as chloride, nitrate, and sulfate out of the root zone. This leaching can result in contamination of groundwater, streams, and lakes or have other environmental implications (Figure 1–20). Excess fertilizer becomes a contaminant and can have adverse effects on human health. The U.S. Environmental Protection Agency has set standards for nutrients in groundwater used for drinking water. This is one of many reasons that appropriate levels of fertilization are essential.

Tips for Collecting a Good Soil Sample

  • Collect samples with stainless steel or chrome-plated tools. Using brass, bronze, or galvanized materials could contaminate the sample.
  • The bucket in which material is collected should be made of plastic.
  • Make sure the collection bucket is clean because even small amounts of residual lime or fertilizer can affect the test results.
  • Avoid taking samples from areas that are obviously different from the norm, such as wet spots, compost piles, animal urine spots, and brush piles, or from under eaves or sites where trash has been burned.
  • Remove large pieces of organic material, such as roots, stalks, and leaves, from the sample.
  • For gardens, new lawns, and other cultivated areas, sample to the depth the soil has been, or will be, tilled. For established lawns, collect the sample 2 to 4 inches deep. For trees and shrubs, take a sample to a depth of 6 inches near the plant’s drip line. Even if the soil looks the same, take separate samples for flower beds, vegetable gardens, fruit orchards, shrub borders, and lawn areas.
  • If using a trowel or spade, dig a hole, then take a slice of soil down one side. Repeat this procedure in five to eight spots for each area to be tested. Mix these cores together to obtain one composite sample. If the soil is very wet, it could be more difficult to mix, but do not attempt to heat the soil to dry it (Figure 1–21).
  • Place about a pint of the composite sample for each area sampled in a soil testing box and label with a return address on the side of the box. Make up a code that will be easy to remember, such as “flawn” for front lawn, “byard” for back yard, or “veg” for vegetable garden. Any combination of letters and numbers can be used. Make notes about where the samples came from so that when you receive the results, you can easily Identify how to treat the areas differently based on the results.
  • Do not tape the boxes in any way. The lids are removed before the boxes go in the soil lab ovens, and tape makes this process difficult. Do not put the soil in a plastic bag before placing it in the box as doing so will prevent proper drying in the lab oven.

Fill out the soil test report sheet, giving as much information as possible. The required items are name, address, county, sample codes, and the crops planned. Reports are sent by mail only if there is a special request submitted to the lab. Otherwise, provide an email address on the form to receive notification that the report is complete and online. Farmers also use the form, so some of the information requested may not apply to gardeners (pounds of lime per acre, for example). Forms and boxes are available from the NCDA & CS or any county Cooperative Extension center.

Learn more about collecting soil samples in SoilFacts: Careful Soil Sampling—The Key to Reliable Soil Test Information (NC Cooperative Extension publication number AG-439-30). For detailed information about the soil test results, refer to NCDA&CS Agronomic Division’s Understanding the Soil Test Report.

How to Use a Soil Test Report

Fertilizing trees and shrubs in a landscape should be based on the amount of rainfall, soil type, the plant’s age, the amount of current growth, and desired future growth. Over application of fertilizer to home landscapes wastes money, contributes to pollution in our rivers, streams, lakes and estuaries, and may damage or kill desired plants. In addition, excess fertilizer can increase the likelihood of disease problems, lead to weak growth, attract pests, and increase the amount of pruning to keep mature plants within appropriate boundaries. A soil test report provides accurate guidance for applying fertilizer.

Example Soil Test Reports

Depending on the crop indicated when the soil sample was submitted, the soil test report provides results in one of two ways:

  • Home garden scale: Recommendations for pounds of lime and a rate and grade of fertilizer per 1,000 square feet (for example, an area 50 feet by 20 feet or 10 feet by 100 feet).
  • Farm/Forest scale: Recommendations for tons of lime and a rate and grade of fertilizer per acre.

At the home gardener scale:

  1. Measure the area to be limed or fertilized.
  2. Multiply the length by the width to determine the number of square feet.
  3. Divide by 1,000 to obtain the number of units to be treated.
  4. Multiply the number of units by the pounds of material to treat 1,000 square feet. This calculation will give the amount of fertilizer and lime needed (Figure 1–23).

Example 1:
If the area is 500 feet by 20 feet, and the suggested lime or fertilizer treatment is 30 pounds per 1,000 square feet:

  1. 500 × 20 = 10,000 square feet
  2. Divide 10,000 by 1,000 = 10 units
  3. Multiply 30 pounds times 10 units = 300 pounds of material (fertilizer or lime) per 10,000 square feet

Example 2:
If the area is 10 feet by 15 feet, and the suggested lime or fertilizer rate is 10 pounds per 1,000 square feet:

  1. 10 × 15 = 150 square feet
  2. Divide 150 by 1,000 = 0.15
  3. Multiply 10 pounds times 0.15 units = 1.5 pounds of material per 150 square feet

Or, look at fertilizer/lime calculations as ratios:

If 5 pounds of fertilizer are applied per 1,000 square feet, how many pounds should be applied to 150 square feet (using the garden size in Example 2)?< /p>

5 lb/1,000 sq ft = X lb/150 sq ft

5 lb × 150 sq ft /1,000 sq ft = X lb

750/1,000 = .75 lb

Example soil test reports and their recommended fertilizer applications can be found in Figure 1–24, Figure 1–25, Figure 1–26, and Figure 1–27.

Soil pH

Soil pH is a measure of the soil’s relative acidity or basicity. The pH scale ranges from 0 to 14. A pH of 7 is a neutral state, representing the value found in pure water. Values above 7.0 are basic, while values below 7.0 are acidic. The pH scale is logarithmic, meaning each unit has a 10-fold increase of acidity or basicity. Thus, compared to a pH of 7.0, a pH of 6.0 is ten times more acidic, and a pH of 5.0 is 100 times more acidic.

Nutrient Availability and pH

The optimum pH for a plant varies with organic matter content and plant type. Plant nutrient availability is strongly tied to the pH in the soil solution (Figure 1–28). Decreasing soil pH directly increases the solubility of the plant nutrients manganese (Mn), zinc (Zn), copper (Cu), and iron (Fe). Acidic soils make these nutrients more available. At pH values less than about 5.5, toxic levels of Mn, Zn, or aluminum (Al), a non-nutrient element very common in our southern soils, may be released. The impact of pH on nutrient availability is very important—both for maximum plant availability and to avoid potentially toxic levels at very low or very high pH.

The optimal pH for growth differs among plants. For example, regardless of organic matter content, azaleas and blueberries are well-suited for a soil pH of about 5.0. In contrast, asparagus can tolerate a basic soil with a pH up to 8.0. A soil pH of 6.5 to 7.0 is often considered “ideal” for most plants, but a little research can help you identify the proper pH for the plants you wish to grow. After obtaining a soil test report, you can take measures to adjust soil pH or select plants that will thrive at the current pH. Extreme pH measures of 4.0 (acidic) or 10.0 (basic) will support little plant life and are very difficult to modify.

Adjusting pH

If the soil pH is too basic for the desired plant, incorporating an acidic soil amendment such as pine bark or compost, or applying elemental sulfur, will lower soil pH. Apply sulfur with caution; too much can harm plants.

If the soil pH is too acidic, apply lime to raise the soil pH. There are two general classes of liming materials: calcitic (without magnesium) and dolomitic (with magnesium). Calcitic lime is composed of calcium carbonate (CaCO3) and can be used on soils high in magnesium. Dolomitic lime is a mixture of calcium and magnesium carbonates (CaCO3 and MgCO3), which is the preferred liming material for soils low in magnesium.

Knowing the soil type or even the current pH is not enough to determine the amount of lime needed. The texture of the soil, organic matter content, crop to be grown, target pH, level of soil acidity, CEC, type and amount of clay, and the current pH are all factors to consider in adjusting pH. Soils low in organic matter or high in sand content require less lime to change the pH than clay soils or those with high organic matter.

Lime is heavily regulated in North Carolina. Lime must be labeled with a guarantee of percent calcium and magnesium. The percent of calcium carbonate equivalent also must be included on the label, as well as the pounds of material that equal 1 ton of standard lime (Figure 1–29). Each type of lime must meet a screening requirement for particle size. Lime pellets are formed from lime that has been finely ground. The pelleted product is less dusty and easier to apply, but is slower to react with the soil.

Lime moves slowly in the soil and neutralizes acidity only in the area where it is applied. To be effective, it should be spread and thoroughly incorporated. It takes several months for lime to react in the soil, which is why it is good to soil test and plan for proper soil pH management. For established lawns, gardens, and ornamentals that require lime, apply the recommended amount up to 50 pounds of lime per 1,000 square feet in one application to the soil’s surface. For recommended rates over 50 pounds, wait several months to make a repeat application to avoid a surface buildup of lime. For new plantings where the area will be tilled, apply the entire recommended amount at one time.

Learn more in SoilFacts: Soil Acidity and Liming: Basic Information for Farmers and Gardeners.


Plant Nutrition and Fertilization

Many people confuse plant nutrition with fertilization. Plant nutrition refers to the needs of the plant and how a plant uses the basic chemical elements. Fertilization is the term used when these elements are supplied to the soil as amendments. Adding fertilizer during unfavorable growing conditions will not enhance plant growth and may actually harm or kill plants.

To complete their life cycle, plants need 17 essential nutrients, each in varying amounts (Table 1–3). Of these nutrients, three are found in air and water: carbon (C), hydrogen (H), and oxygen (O). Combined, C, H, and O account for about 94% of a plant’s weight. The other 6% of a plant’s weight includes the remaining 14 nutrients, all of which must come from the soil. Of these, nitrogen (N), phosphorus (P), and potassium (K), the primary macronutrients, are the most needed. Magnesium (Mg), calcium (Ca), and sulfur (S), the secondary macronutrients, are next in the amount needed. The eight other elements—boron, chlorine, copper, iron, manganese, molybdenum, nickel, and zinc—are called micronutrients because they are needed in much smaller amounts than the macronutrients.

Table 1–3. Relative Amounts (out of 100) of the Essential Nutrients Required by Most Plants

Primary Nutrients

Carbon (C)45
Oxygen (O)45
Hydrogen (H)6
Nitrogen (N)1.5
Potassium (K)1
Phosphorus (P)0.2

Secondary Nutrients

Calcium (Ca)0.5
Magnesium (Mg)0.2
Sulfur (S)0.1


Iron (Fe)0.01
Chlorine (Cl)0.01
Manganese (Mn)0.005
Boron (B)0.002
Zinc (Zn)0.002
Copper (Cu)0.0006
Molybdenum (Mo)0.00001
Amounts unknown for Nickel (Ni) and Cobalt (Co)

Soil Nutrients

For a plant to absorb an element, it must be in a chemical form used by the plant and dissolved in the soil water. In addition to those nutrients already dissolved in soil water, nutrients can be present in the soil in these forms:

  • Undissolved or granular form, as from newly applied fertilizer
  • Chemicals bound to soil particles
  • The chemical structure of soil organic matter released by microbial decomposition

Undissolved or granular nutrients, and those that are chemically bound to soil particles, are not immediately useful, although they have the potential to benefit the plant. For many plant nutrients, the soil acts as a bank. Withdrawals are made from the soil solution, much as you would withdraw money from a checking account. The undissolved pool of soil nutrients is like a savings account. When checking funds are low, transfers are made from the savings account to the checking account. When a checking account is flush with money, some can be moved to savings for long-term retention. In the same way, for many plant nutrients, when the soil solution has excess nutrients, some bind to the soil to become temporarily unavailable, and some react with other chemical elements to form insoluble minerals, which can dissolve again later.

Several factors improve a plant’s ability to use nutrients:

  • Type of soil: The more clay and organic matter a soil has, the higher its CEC will be, and the more cationic (positively charged) nutrients it will retain.
  • Soil pH: The pH affects how tightly nutrients are bound to soil particles. If the soil pH is extremely high (basic) or very low (acidic), many nutrients become inaccessible to the plant because they are no longer dissolved in the soil water.
  • Types of nutrients in the soil: Some nutrients affect the availability of other nutrients. In fact, an apparent deficiency of one nutrient may actually be caused by a large amount of another.
  • Amount of soil water: Too much rain leaches nutrients from the soil. If there is too little water, the nutrients cannot dissolve and move into the plant.
  • Anything that affects the plant’s growth: If growing conditions are good, a plant will absorb nutrients from the soil. If the plant experiences extremes in temperature, incorrect light levels, or waterlogged or compacted soil, it will have a limited ability to absorb nutrients. Also, plants in dormant stages absorb few nutrients.

The presence or absence of nutrients can cause outward symptoms to appear on the plant. Table 1–4 reviews the essential nutrients for plant growth and symptoms that may appear if a plant is suffering a deficency or an excess of that nutrient.


Fertilizers provide some elements that might be lacking in the soil and stimulate healthy, vigorous growth. How much and when to apply fertilizers should be based on observing plant performance, a reliable soil test, and an understanding of the factors that affect growth: light, water, temperature, pests, and nutrition. Simply applying fertilizer because a plant is not growing adequately will not solve many plant problems (insects, disease, or poor drainage, for example), and, in fact, excess nitrogen can often increase insect and disease infestation.

All fertilizers are labeled with three numbers, giving the percentage (by weight) of nitrogen (N), phosphorus (P), and potassium (K). This is referred to as the fertilizer grade.

A 100-pound bag of fertilizer labeled 0-20-10 has 0 pounds of N, 20 pounds of P (reported as P2O5), 10 pounds of K (reported as K2O), and 70 pounds of filler. Filler is added to make the fertilizer easier to spread and to reduce the likelihood of burning plants with too much fertilizer (the fertilizer salts can pull water out of the plant). A fertilizer may also contain secondary macronutrients or micronutrients not listed on the label because the manufacturer does not want to guarantee their exact amounts.

Fertilizers can be divided into two broad categories: natural and synthetic.

Natural fertilizers are commonly misnamed “organic.” “Natural fertilizers” is a more accurate description because these materials can be both complex chemical substances containing carbon (organic materials) or inorganic ores, such as rock phosphate, which are mined. Natural fertilizers containing organic materials include manures and composts, animal byproducts (such as bone meal, blood meal, feather meal), and seed meals. Natural fertilizers that are inorganic ores include potassium and lime.

Natural fertilizers typically release nutrients at a slower rate and over a longer period than synthetic fertilizers because microorganisms are involved in a breakdown and release cycle called mineralization. Moisture, temperature, and the microbial species and populations in the soil affect mineralization. Some water-soluble natural fertilizers, such as fish emulsion, are available when rapid nutrient delivery is desired.

When using natural fertilizers, it is helpful to incorporate them and provide adequate moisture for active microbial populations. When packaged as fertilizers, natural fertilizers will have the nutrient analysis stated on the labels. How much to use varies with the nutrient content of the material. The age of the material is also a factor. Producers are not required by law to state the nutrient content on bulk organic materials, such as compost, manure, and sludges. The source of these materials should be investigated and possible analysis performed at the Plant, Waste, Solution, and Media Lab at the NCDA&CS Agronomic Division before applying large amounts to a home garden.

The age of the natural fertilizers is another important factor. When natural material decays and is rained on, it loses nutrients, especially potassium and, to some extent, nitrogen. Even natural sources of nutrients can be overappled and damage plants. Fresh manures, for example, may injure plants by adding excessive nitrogen or potassium, especially when applied in large quantities.

Natural fertilizers can be expensive if applied in amounts adequate to supply nutrients for good plant growth, but have the added benefit of improving soil structure and plant vigor. When applying natural fertilizers, calculate as closely as possible the amounts of nutrients being supplied. Always err on the low side of application rates, then test the soil and augment as recommended on the soil test report. The nutrient content may need to be supplemented with other natural or synthetic materials to achieve a balanced ration of nutrients.

Synthetic fertilizers are made through industrial processes or mined from deposits in the earth. They are purified, mixed, blended, and altered for easy handling and application. Most are noncarbonaceous chemicals from nonliving sources and are usually cheaper than natural fertilizers. In general, nutrients are more rapidly available to plants because they are more water-soluble or in a form plants can use. The disadvantage is that it may be easier to over apply a synthetic fertilizer than a natural one, which may result in fertilizer burn. In addition, synthetic fertilizers may not support beneficial microbial populations to the same extent as natural fertilizers.

Special-purpose fertilizers are packaged for plants such as camellias, rhododendrons, and azaleas (Figure 1–43). Some of the compounds used in these fertilizers have an acid reaction that can be beneficial to acid-loving plants if the soil they are growing in is naturally neutral or alkaline; however, most soils in North Carolina are usually acidic so these special fertilizers are unnecessary.

Fertilizer spikes or pellets are fertilizers compressed into a form placed in the soil or pots (Figure1–44). They are convenient, but are expensive per unit of fertilizer and do not provide uniform distribution. Nutrients are often concentrated around the spikes or pellets.

Liquid fertilizer can be purchased as a dry powder or as a concentrated liquid (Figure 1–45). Liquid fertilizers are frequently used for houseplants or as a starter solution for transplants. They tend to be more expensive per unit of fertilizer because they are made from refined chemicals.

Foliar fertilizers are dry powders or concentrated liquids that are mixed with water and sprayed on plants (Figure 1–46). Foliar feeding is used when insufficient fertilizer was applied before planting, when a quick growth response is wanted, when micronutrients are locked in the soil, or when the soil is too cold for the plant to use fertilizer in the soil. Foliar-applied nutrients are absorbed and used by the plant quite rapidly. They are expensive per unit of nutrient and only give short-term fertilization (completely absorbed within one to two days). Relying totally on foliar fertilization can be time consuming because the fertilizer must be applied regularly. Improper foliar application of fertilizers can also lead to plant tissue burn.

Learn more about fertilizer usage and nutrient concentrations in the North Carolina Agricultural Chemicals Manual, Chapter IV – Fertilizer Use.

Fertilizer Terms

Fertilizer analysis: The minimum amount of each element in a fertilizer as stated on the label, such as 16-4-8.

Fertilizer ratio: The relative proportion of N, P2O5, and K2O. The ratios of 16-4-8 and 8-2-4 are both 4:1:2, which means 4 parts nitrogen to 1 part phosphorus to 2 parts potassium.

Balanced fertilizer: A fertilizer containing equal parts of each major element, such as 10-10-10.

Complete fertilizer: A fertilizer containing nitrogen, phosphorus, and potassium. Examples of commonly used fertilizers are 10-10-10, 16-4-8, and 12-4-8.

Incomplete fertilizer: A fertilizer missing one or two of the macronutrients, such as 0-20-0.

Weed and feed fertilizers: A combination of fertilizer and herbicide. They are often used on lawns to prevent certain weeds from germinating, or to kill existing broadleaf weeds.

High analysis: A fertilizer containing 30% or more active nutrients, such as ammonium nitrate 33-0-0. The cost per bag is usually more, but the cost per pound of nutrient is less, lowering the cost for fertilizing a given area.

Incomplete fertilizers can be used separately or combined to supply the needed nutrients, often at a reduced cost compared to using a complete fertilizer. For example, gardeners who have a soil with sufficient P and K can save money by applying a nitrogen-only fertilizer, such as ammonium nitrate (34-0-0). If a soil test indicates N and K are needed, but not P, use an appropriate amount of ammonium nitrate and muriate of potash (0-0-60), a naturally occurring material composed almost entirely of potassium, processed to remove impurities and concentrate the product. If a soil needs only P, use triple super phosphate (0-46-0), or for an organic nutrient source apply bone meal (approximately 3-15-0; note that this will add some N) or compost.

Regardless of the fertilizers used, be aware that excess fertilizer can damage plants and move into our stormwater systems, which can cause serious environmental problems.

Plant Nutrients and the Environment

Fertilizer misuse causes environmental and water quality issues. Nitrogen fertilizers, for instance, break down into ammonium and nitrate. The nitrate form of N, while essential for plant growth, is highly mobile and can move through the soil after rainfall or irrigation and contaminate drinking water supplies. Phosphorus holds tightly to soil particles and does not leach through the soil, but affects water quality through runoff and soil erosion. Excess nitrogen and phosphorus are associated with algal blooms (heavy growth of aquatic plants) and limited oxygen, and cause fish kills in lakes, bays, and nonflowing water bodies.

There are several ways to reduce fertilizers’ impacts on water quality:

  • Apply only materials that are recommended based on results of a soil test. If possible, use slow-release fertilizers and incorporate into the soil. Avoid applying excess nitrogen and phosphorus fertilizer.
  • Calibrate fertilizer spreaders properly and clean spreaders over the lawn area instead of a hard surface.
  • Keep the amount of hard surfaces in a landscape to a minimum. When installing a new sidewalk or patio, consider using gravel, porous concrete, stepping-stones, wood decking, or bricks on a sand base (Figure 1–47).
  • Avoid applying fertilizer to hard surfaces, such as sidewalks, patios, driveways, and streets. Sweep up material that falls on hard surfaces.
  • Maximize water absorption by aerating lawns and incorporating organic matter in planting beds and gardens.
  • Prevent runoff by turning off irrigation when the soil is no longer absorbing water.
  • Avoid applying fertilizer in natural drainage areas or ditches.
  • Minimize soil erosion by using ground covers, windbreaks, terraces, and mulches.
  • Mulch under trees and shrubs to reduce impact of falling water.
  • Maintain a lawn border around planting areas and plant a grass strip between rows in fruit and vegetable gardens.
  • Plant cover crops on bare soil, such as barren vegetable gardens.
  • Use a rain barrel under drains to collect runoff and direct excess runoff from roofs onto grassy areas (Figure 1–48).

When to Apply Fertilizer

Soil type affects the frequency of fertilizer application. Sandy soils require more frequent applications of smaller amounts of nitrogen and potash than do clay soils because these nutrients leach more readily in sandy soils. Other factors that affect application frequency include the plant to be grown, the amount of plant growth desired, the amount of water, and the type and release rate of fertilizer applied.

The best time to apply fertilizer and the most effective method of applying it depend on the type of plants being grown. Leafy vegetables require more nitrogen than root crops. Corn is a heavy nitrogen feeder and may require several small nitrogen applications when actively growing. Most established woody plants perform well without fertilization, or with just one application per year. Young plants may benefit from several light applications of fertilizer per year.

Fertilizer is needed when plants are actively growing, never when they are dormant. Nitrogen application will have its greatest effect three to four weeks after application. Excess or improperly timed nitrogen can delay flowering and fruiting or promote tender new growth vulnerable to frost or freeze damage.

Research has shown it is best to broadcast or incorporate fertilizer uniformly over an area rather than concentrating fertilizer in holes or bands in the soil. The most effective method of fertilizing a large area is with a fertilizer spreader; for home gardens, hand fertilization works fine. For new plantings, incorporate fertilizer into the soil and mix it thoroughly. For established plantings, surface application is appropriate.

When fertilizing from overhead, make certain plant foliage is dry and use a broom to brush fertilizer off the foliage, or water thoroughly after applying fertilizer to remove it from plant leaves to prevent burn spots. It is not necessary to remove mulch when fertilizing; irrigation or rainfall will carry fertilizer to the roots. Fertilization should be reduced or delayed during dry weather because the salts in the fertilizer can burn roots if there is inadequate moisture.

Calibrating a Spreader

Fertilizers are more effective if they are applied at the proper rate and with uniform coverage. To accomplish this, calibrate the spreader, which requires a little labor and math.

The two types of spreaders used to apply fertilizer and lime are drop spreaders (Figure 1–49) and rotary spreaders (Figure 1–50). The amount of fertilizer that is spread depends on the opening setting, the type of fertilizer, and the speed at which the spreader is pushed. The drop spreader has a series of holes at the base that can be adjusted to apply different amounts of material. With the rotary spreader, the fertilizer falls into a rotating plate and is spread by the centrifugal force of the plate spinning. Instructions for calibrating a spreader should be available on the Internet at the home page for the spreader manufacturer.

The Biology of Soil

There is more life below the soil surface than there is above. Soil life consists of burrowing animals, such as moles and earthworms, insects, and other soil creatures that are difficult or impossible to see without a microscope, such as mites, springtails, nematodes, viruses, algae, bacteria, yeast, actinomycetes, fungi, and protozoa. There are about 50 billion microbes in 1 tablespoon of soil. In a typical soil, each gram (what a standard paperclip weighs) likely contains these organisms, listed from largest to smallest:


  • Nematodes—10 to 5,000
  • Algae—1,000 to 500,000
  • Protozoa—1,000 to 500,000
  • Fungi—5,000 to 1,000,000
  • Actinomycetes—1,000,000 to 20,000,000
  • Bacteria—3,000,000 to 500,000,000

Soil-dwellers move through the soil, creating channels that improve aeration and drainage. Nematodes and protozoa swim in the film of water around soil particles and feed on bacteria. Mites eat fungi, and fungi decompose soil organic matter. The microorganisms’ primary role is to break down organic matter to obtain energy. Microorganisms help release essential nutrients and carbon dioxide and perform key roles in nitrogen fixation, the nitrogen and phosphorus cycles, denitrification, immobilization, and mineralization. Microbes must have a constant supply of organic matter, or their numbers will decline. Conditions that favor soil life also promote plant growth.


Unfavorable soil conditions, such as high temperatures, compaction, or oversaturation can injure beneficial soil life. This can lead to a proliferation of disease-causing fungi, bacteria, or viruses. To read more about common soil diseases see chapter 5,“Diseases and Disorders.” Plants that are stressed by disease are often more susceptible to insect damage. More information on insects can be found in chapter 4, “Insects.” To learn more about managing insects and diseases, please see chapter 8, “IPM .”


To promote soil organisms, incorporate organic matter, till as little as possible, minimize soil compaction, maintain favorable soil pH and fertility, and use organic mulch on the soil surface.

Further Reading


Brady, Nyle C., and Ray R. Weil. The Nature and Properties of Soils. 14th ed. Upper Saddle River, New Jersey: Prentice Hall, Inc, 2007. Print.


Buol, S. W., et al. Soil Genesis and Classification. 6th ed. Hoboken, New Jersey: John Wiley & Sons Inc., 2011. Print.


Dunne, Niall, ed. Healthy Soils for Sustainable Gardens. Brooklyn, New York: Brooklyn Botanic Garden, 2009. Print.


Maynard, Donald N., and George J. Hochmuth. Knott’s Handbook for Vegetable Growers. 5th ed. Hoboken, New Jersey: John Wiley & Sons, Inc., 2007. Print.


Soil Fertility Manual. 5th ed. Peachtree Corners, Georgia: International Plant Nutrition Institute, 2003. Print.


Table 1–4. Essential Nutrients for Plant Growth

NutrientWhy Nutrient is NeededDeficiency SymptomsExcess SymptomsComments
Nitrogen (N)• Responsible for rapid foliage growth and green color
• Easily leaches from soil
• Mobile in plant, moving to new growth
• Reduced growth (Figure 1–30)
• Light-green to yellow foliage (chlorosis)
• Reds and purples may intensify with some plants
• Reduced lateral breaks
• Symptoms appear first on older growth
• Succulent growth; leaves are dark green, thick, and brittle
• Poor fruit set
• Excess ammonia can induce calcium deficiency
• High N under low light can cause leaf curl
• Uptake inhibited by high P levels
Phosphorus (P)• Promotes root formation and growth
• Affects quality of seed, fruit, and flower production
• Increased disease resistance
• Does not leach from soil readily
• Mobile in plant, moving to new growth.
• Reduced growth
• Leaves dark-green; purple or red color in older leaves, especially on the underside of the leaf along the veins (Figure 1–31)
• Leaf shape may be distorted
• Thin stems
• Limited root growth
Shows up as micronutrient deficiency of Zn, Fe, or Co• Rapidly fixed on soil particles
• When applied under acid conditions, fixed with Fe, Mn, and Al
• High P interferes with micronutrient and N absorption
• Used in relatively small amounts when compared to N and K
• Availability is lowest in cold soils
Potassium (K)• Helps plants overcome drought stress
• Improves winter hardiness
• Increases disease resistance
• Improves the rigidity of stalks
• Leaches from soil
• Mobile in plant
• Reduced growth
• Shortened internodes
• Margins of older leaves become chlorotic and burned
• Necrotic (dead) spots on older leaves (Figure 1–32)
• Reduction of lateral breaks and tendency to wilt readily
• Poorly developed root systems
• Weak stalks
Causes N deficiency and may affect the uptake of other nutrients• High N/low K favors vegetative growth
• Low N/high K promotes reproductive growth (flower, fruit)
• Calcium excess impedes uptake of K
Magnesium (Mg)• Leaches from sandy soil
• Mobile in plant
• Reduction in growth
• Yellowish, bronze, or reddish color of older leaves, while veins remain green (Figure 1–33)
• Leaf margins may curl downward or upward with a puckering effect
• Interferes with Ca uptake
• Small necrotic spots in older leaves
• Smaller veins in older leaves may turn brown
• In advanced stage, young leaves may be spotted
• Mg is commonly deficient in foliage plants because it is leached and not replaced
• Epsom salts at a rate of 1 teaspoon per gallon may be used two times a year
• Mg can be absorbed by leaves if sprayed in a weak solution
• Dolomitic limestone can be applied in outdoor situations to rectify a deficiency
Calcium (Ca)• Moderately leachable
• Limited mobility in plant
• Essential for growth of shoot and root tips
• Inhibition of bud growth
• Roots can turn black and rot
• Young leaves are scalloped and abnormally green
• Leaf tips may stick together
• Cupping of maturing leaves
• Blossom end rot of many fruits (Figure 1–34)
• Pits on root vegetables; stem structure is weak
• Premature shedding of fruit and buds
• Interferes with Mg absorption
• High Ca usually causes high pH
Ca is rarely deficient if the correct pH is maintained
Sulfur (S)• Leachable
• Not mobile
• Contributes to odor and taste of some vegetables
• Rarely deficient
• General yellowing of the young leaves, then the entire plant (Figure 1–35)
• Veins lighter in color than adjoining interveinal area
• Roots and stems are small, hard, and woody
Sulfur excess is usually in the form of air pollutionSulfur excess is difficult to control, but rarely a problem


NutrientWhy Nutrient is NeededDeficiency SymptomsExcess SymptomsComments
Iron (Fe)• Accumulates in the oldest leaves and is relatively immobile
• Necessary for the maintenance of chlorophyll
• Interveinal chlorosis primarily on young tissue, which may become white (Figure 1–36)
• Fe deficiency may occur even if Fe is in the soil when: soil is high in Ca; soil is poorly drained; soil is oxygen deficient; nematodes attack roots; or soil is high in Mn, pH, or P
• Fe should be added in the chelate form; the type of chelate needed depends upon the soil pH
• Foliar fertilization will temporarily correct the deficiency
• May be deficient in centipede grass where pH and P are high
Rare except on flooded soils 
Boron (B)• Important in enabling photosynthetic transfer
• Very immobile in plants
• Failure to set seed
• Internal breakdown of fruit or vegetable
• Death of apical buds, giving rise to witches’ broom
• Failure of root tip to elongate normally
• Young leaves become thick, leathery, and chlorotic (Figure 1–37)
• Rust-colored cracks and corking on young stems, petioles, and flower stalks (such as heart rot of beets, stern crack of celery)
• Breakdown occurs at the base of the youngest shoots

• Tips and edges of leaves exhibit necrotic spots coalescing into a marginal scorch (similar to high-soluble salts) (Figure 1–38)
• Oldest leaves are affected first • Can occur in low pH soils • Plants are easily damaged by excess application •Looks like Mg deficiency,green veins on a yellow leaf.

Zinc (Zn)Needed for enzyme activity• Young leaves are very small, sometimes missing leaf blades
• Short internodes
• Distorted or puckered leaf margins
• Interveinal chlorosis (Figure 1–39)
• Severe stunting, reddening
• Poor germination
• Older leaves wilt
• Entire leaf is affected by chlorosis; edges and main vein often retain more color
• Can be caused by galvanized metal.
Copper (Cu)Needed for enzyme activity• New growth small, misshapen, wilted (Figure 1–40)
• In some species, young leaves may show interveinal chlorosis while tips of older leaves remain green
• Can occur at low pH
• Shows up as Fe deficiency
Manganese (Mn)Needed for enzyme activity• Interveinal chlorosis with smallest leaves remaining green, producing a checkered effect (Figure 1–41)
• Grey or tan spots usually develop in chlorotic areas
• Dead spots may drop out of the leaf
• Poor bloom size and color
• Induced by excessively high pH
• Reduction in growth, brown spotting on leaves
• Shows up as Fe deficiency
• Found under strongly acidic conditions
Molybdenum (Mo)Needed for enzyme activity• Interveinal chlorosis on older or midstem leaves (Figure 1–42)
• Twisted leaves whiptail
• Marginal scorching and rolling or cupping of leaves
• Nitrogen deficiency symptoms may develop
• Intense yellow or purple color in leaves
• Rarely observed
Chlorine (Cl)Needed for enzyme activity• Wilted leaves which become bronze, then chlorotic, then die
• Club roots
• Salt injury
• Leaf burn
• May increase succulence
Cobalt (Co)• Needed by plants recently established
• Essential for nitrogen fixation
Little is known about its deficiency symptomsLittle is known about its toxicity symptoms 
Nickel (Ni)• Needed by plants recently established
• Essential for seed development
Little is known about its deficiency symptomsLittle is known about its toxicity symptoms 


Potting Soil

The structure of potting soil determines how much water and air are available to the roots. Air is important for the roots’ oxygen supply and for the micro-organisms in the root environment.


infopaper-terra_text_4.jpgRoots need oxygen for growth, maintenance of the root system, and the uptake of water and nutrients. The provision of a good supply of air for fast growing plants is of vital importance for obtaining good results. A shortage of air in the early stages of cultivation leads to a poorly developed root system, which hampers growth resulting in a smaller yield.


Air (Oxygen) moves from the air supply outside or in the growing room, through large pores in the medium to the root surface by diffusion. The structure of the potting soil in use is dependent on the quality of the raw materials comprising the soiless potting mix. The best soiless potting media starts with virgin peat that has long-term stable structural characteristics.


Vertically cut or shaved peat?


infopaper-terra_text_5.jpgTwo different methods of harvesting can be used to exploit peat deposits: the cheapest but least efficient method consists of “shaving off” the top layer of peat. The disadvantage of this technique is that the structure is less coarse which has an undesirable effect on the air/ water relationship.


The second method consists of cutting the peat vertically. This is the most well known method from times past. The peat blocks that were previously used as fuel for cooking and heating houses were cut in this way. This is an expensive way of extracting peat. When peat is cut into blocks it has to be turned by hand to dry out.


The coarser the peat is, the better will be the balance between the water and air it contains. This ensures the roots develop better in the medium. The plants are healthier and the tendency of the mix to compress is reduced.


High and low peat


One of a potting medium’s main ingredients is peat. This is a century’s old, naturally occurring material formed from old vegetation. Peat originates from regions where climatic circumstances caused new plant material to form faster than the dead vegetation could rot. Slowly but surely this process built up a layers of organic materials that become several meters deep in some locations.


Two different types of peat can be differentiated: high peat and low peat. Low peat is formed in regions where groundwater levels are high and there are plenty of nutrients. Low peat typically contains a high proportion of rotten material and can have high levels of silt; sand and harmful salts which makes it an unsuitable base for a good potting soil. High peat is formed under the influence of rainwater in low nutrient regions and its principal ingredient is sphagnum moss. This is a small plant that dies off from below while continuing to grow above on its own remains. The remains of other plants are also found here; grasses, tree limbs and leaves for example.



Sphagnum moss


Sphagnum moss is particularly suitable for use as a growing substrate because it is natural, light, clean and easy to work with. It has a low nutritional content; a pH that ranges from 3.5 to 4.5 and has good water retention properties (up to 20 times its own weight). The profile of high peat in the flat peat region of Northwest Germany, contains different, distinct layers that formed in different periods.

Micro-organisms in the growing medium

Micro-organisms are present everywhere; in the air, in water, on plants and in the soil. They can remain dormant for long periods in many different ways, as spores, mitochondrion (a membrane-enclosed organelle found in the cells of most organisms) or hyphae, and they can live up to several years and on a wide range of hosts. Although most people think of micro-organisms as being harmful by definition, life as we know it would not be possible without these minuscule life forms. In this article we will look at how they affect the growth of plants through their presence in the growing medium. By CANNA Research


Micro-organisms include a very broad range of species, and include bacteria, protozoa, algae and fungi. Most of these micro-organisms can multiply rapidly when the circumstances are right and they can have a major influence, both positive and negative, on the development of plants growing in the substrate. The kinds of micro- organism that appear in a given substrate will depend on a number of factors, such as the climate, the properties of the substrate, the plant species and the other organisms present in the substrate.


Micro-organisms in the growing medium

Micro-organisms can significantly affect the development of plants growing in the substrate, both positively and negatively. Many factors are important for micro-organisms, such as the type of growing medium. Others include the amount of oxygen, the temperature of the substrate and roots, the pH level and the salinity or EC level in the substrate. Another aspect that has quite an impact on the micro-life in the growth medium is the presence of pesticides.


Peat, coco coir, rock wool or perlite


One of the most important factors that influence micro-organisms is the type of growing medium available to them. The main differences can be found between organic substrates, such as coco coir, peat or soil, and inorganic substrates such as clay pebbles, perlite or rock wool. Micro-organisms that have a high saprophytic capacity (i.e. they live off dead organic matter) will do better in a substrate containing organic material such as peat or re-used substrate. But dead leaves lying on the surface of the substrate or infected plants with necrotic parts will do just as well.


The amount of oxygen in the substrate determines if aerobic or anaerobic micro-organisms will develop. Most plants need oxygen around their roots, so normally aerobic organisms will be present. Plants grown in a substrate with very little oxygen will usually be weaker and pathogenic anaerobic micro-organisms can benefit from that.


All organisms have an optimum growing temperature, so the temperature of the substrate and roots will affect the micro-flora around them. The average temperature and the temperature range (cold nights or a hot summer day) will determine which micro-organisms can survive, as well as the range and frequency of temperature fluctuations.


Micro-organisms in the growing medium
Fungus growth on rock wool cubes.


Just like plants, most micro-organisms prefer an acidity level of between pH 5.5 and 5.8. high and low pH levels, as well as pH fluctuations can disturb the development of micro-flora. still, some micro-organisms are able to grow or even flourish in extreme conditions. The salinity or EC level in the substrate will also affect the growth of the micro-organisms, and the type and composition of the salts in the growing medium also has an impact. This is especially true of salts such as Potassium chloride or Sodium chloride that can change the rhizosphere of plants and thereby the kinds of micro-life which will populate the root zone.


In inorganic substrates such as rock wool, clay pebbles or perlite, most micro-life is waterborne. These micro-organisms are introduced via watering systems, air or plants and will survive as long as the moisture level is sufficiently high, even if there are only small pockets of moisture. As with soil-borne micro-organisms, these can also be either beneficial or pathogenic.


Another aspect that has quite an impact on the micro-life in the growth medium is the presence of pesticides. These can build up in substrates or soils that are used for a prolonged period of time. Depending on the kind of pesticides – either herbicides, insecticides, fungicides or bactericides – these will influence the composition of microscopic life in the substrate.


One thing that one can rely on is the fact that micro-organisms can adapt to a range of circumstances. A well-known example is the resistance of some bacteria against antibiotics; it only takes a fractional change in their genetic material, but the effect on the resistance can be all-important.


The benefits of micro-organisms


The presence of micro-organisms can have both a positive and negative impact. As such, it is not necessary or desirable to get rid of all micro-organisms. The ability of a crop to defend itself against infections depends largely (albeit not exclusively) on the presence of micro-life in the substrate. It is difficult to quantify this benefit, however, since there are numerous factors on which the micro-life depends. The defense relates to the total microbial activity, the diversity of different groups of functional actinomycetes (rod-shaped bacteria), the total population of actinomycetes and the percentage of cellulose-decomposing actinomycetes.


If there is a good balance of micro-life in the substrate, there will most likely be less need to use pesticides and other measures, which will reduce costs. Not only are fewer and fewer pesticides actually permitted in horticulture and other applications, they are also very expensive. Plus, in a sterile substrate the most opportunistic micro-organisms will find a free space with no competitors and unlimited access to space and nutrients. These first opportunistic colonists will not necessarily be beneficial to the crop planted in the substrate. It is wiser to use the correct micro-life from the beginning to produce a healthy crop and good yield. Micro-organisms can even be used to improve the quality of reused substrates, where certain bacteria actually produce enzymes that can decompose accumulated salts.


Micro-organisms in the growing mediumPythium infection on tomato roots in coco coir.


Substances that are exuded from the root system like sugars, amino acids or phenols can either attract or repel micro-organisms. The position of each species in the competition game that is continually going on between the bacteria can be influenced by these root exudates. Plants can use this to its advantage by exuding substances that attract beneficial organisms, establishing a symbiosis with a particular micro-organism. A prime example is the presence of symbiotic bacteria in the root nodules of leguminous plants such as peas or beans, which convert atmospheric nitrogen into a form that can be absorbed and used by the plant.


Another tactic is to introduce certain benign organisms to suppress pathogens. This works as follows. some micro-organisms are not very competitive and have a hard time colonizing a substrate that is already occupied by other micro-organisms. This can serve as a mechanism to get rid of pathogenic micro-organisms. By introducing beneficial micro-organisms such as mycorrhiza or trichoderma fungi into a clean substrate, the growing medium will become less inviting for pathogenic micro-organisms, thus protecting the plant from becoming infected.


Beneficial microbes compete with pathogens for nutrients or glucose and some antagonists have their own method of winning a competitive edge. For example, the fluorescent Pseudomonas bacteria can produce proteins that transform slightly soluble iron (Fe) into iron chelate, which it can then absorb much more easily. This then deprives the Fusarium fungi of the iron it needs to grow, preventing it from developing. competition for glucose can also cause microbiostasis which means that the spores of that pathogenic fungus germinate much more slowly due to a lack of energy from glucose.


Antagonistic micro-organisms can also block one or more stages of the propagation cycle of pathogens. Pseudomonas species P. Stutzeri, for example, interrupts the formation of conidia (asexual spores of several kinds of fungi) and the formation and germination of chlamydospores (thick-walled dormant spore of several kinds of fungi), but has no effect on mycelial growth (mycelial cords are capable of transferring nutrients over long distances). Pseudomonas can also produce antibiotics, which can be another tactic to remove pathogens, while other micro-organisms produce enzymes that attack the cell walls of competing species. Antagonists that produce chitinolytic enzymes have the potential to act against pathogenic fungi. It has also been found that several antagonistic organisms or closely related species can co-operate to fight a pathogen. Other antagonists simply overwhelm a pathogenic micro-organism by multiplying more rapidly and thus depriving all the competition of resources and therefore any chance of survival.


Micro-organisms in the growing medium
Algae development on rock wool cubes.


Pathogenic or harmful micro-organisms


Micro-life in the substrate also comes in the form of soil- or water-borne pathogens. some of these pathogens can attack over 80 different plant species and their resilience means they can be very important. There are many different harmful micro-organisms which result in a range of infections and symptoms (rotting fruits, fading, and necrosis to name but three).


Some pathogens produce micro-toxins which can attack the plant or the micro-life in the substrate. Pathogens can gain an advantage over other micro-organisms when they are able to germinate faster and are able to remain dormant for a longer period when conditions are not optimal. Pathogenic and non-pathogenic species can be very closely related, which makes it hard to use antagonists or other measures. Pathogens can fight back when they are attacked by antagonists. an example is fusarium, which can produce fusarium acid that affects the plant cells but can also suppress the production of antibiotics of Pseudomonas (this was discovered through research at Wageningen University in the Netherlands).


Balance your micro-organisms


Most infections by pathogens are in fact the result of a plant that was weak to begin with. Healthy plants are resilient and will be able to respond to an infection by micro-organisms. Provided the plant’s responses are quick and strong enough, it will be able to overcome such an infection. As such, ensuring an optimal climate and soil conditions for the crop is even more important than optimal growing conditions for (beneficial) micro-organisms.


It is crucial to get a good balance of micro-organisms in the substrate over a prolonged period of time, yet sometimes the growth cycle of a plant is simply too short to achieve this balance. Inoculating the substrate with antagonists is possible and, although the results are not always consistent, in some cases this can have a very positive effect on plant growth and health. In some cases, the results are comparable to the effect of using chemicals like fungicides, although these results may not last throughout the entire growing season.


Micro-organisms in the substrate can be a great help in suppressing plant diseases and a great deal of research is being conducted in this area of horticulture. Although this technology has still not entered the mainstream, research by the Louis Bolk Institute in the Netherlands has shown that introducing beneficial micro-organisms and or adding compost to increase the amount of micro-life can have a major effect on crop performance.